ХронологияБиблиотекаMesolithic Western Eurasian DNA

Сообщение Опубликовано » 13 сен 2015, 17:07, последние изменения » 14 янв 2016, 18:09





Mesolithic Western Eurasian DNA

Mitochondrial and Y-chromosome haplogroups extracted from prehistoric hunter-gathers after the Last Glacial Maximum in Europe and related remains in Asia, arranged chronologically. Pitted Ware sites are classed as Mesolithic here, because of their hunter-gatherer economy, though they fall within the Neolithic period.

Culture Country Site and/or Individual Sex Date Y-DNA mtDNA Source
    pigment. from genes   usually approx. Haplogroup Additional information Haplogroup Additional information  
  England, UK Gough's Cave, Cheddar [Not Cheddar Man]   12,700 BC     Unreliable CRS. Contamination likely. Sykes 2000; Jacobi 2009 for dating
  Italy Villabruna, Trentino-South Tyrol   12,267-11,831 BC     ? 16261, 16274 Di Benedetto 2000
  Portugal           Unreliable 16336, reported as H. Museum bones, contamination likely. Chandler 2005
  Portugal           Unreliable ; U5b1c1? 16311, 16336, reported as H. Museum bones, contamination likely. Chandler 2005
  Portugal           Unreliable; V10a? 16222, 16261, 16311, reported as H. Museum bones, contamination likely. Chandler 2005
  Portugal           Unreliable; U4/H1b? 2 samples 16189, 16356, 16362, reported as H. Museum bones, contamination likely. Chandler 2005
  Portugal           Unreliable; N5? 16073, 16111, 16223, reported as N*. Museum bones, contamination likely. Chandler 2005
  Portugal           Unreliable; N1b? 16073, 16223, 16390, reported as N*. Museum bones, contamination likely. Chandler 2005
  Portugal           Unreliable; U4 16073, 16189, 16356, 16362, reported as U* Chandler 2005
  Portugal           Unreliable; U5b1c2 16073, 16174, 16189, 16192, 16270, 16311, reported as U5. Museum bones, contamination likely. Chandler 2005
  Germany Blätterhöhle, Hagen [BLA 3]   9210 ± 29 BC     U2e 16129C, 16189C, 16362C, 16391A Bollongino 2013
  Germany Blätterhöhle [BLA 6]   8796 ± 90 BC     U5b2a2 16172C, 16189C, 16270T, 16398A Bollongino 2013
  Germany Blätterhöhle [BLA 2]   8748 ± 67 BC     U/K 12308G Bollongino 2013
  Germany Blätterhöhle [BLA 20]   8652 ± 58 BC     U5a2c3* 73G, 263G, 750G, 1438G, 2706G, 3197C, 4769G, 7028T, 8860G, 9477A, 10619T, 11465C, 11467G, 11719A, 12308G, 12372A, 13617C, 14766T, 14793G, 15326G, 16192T, 16256T, 16270T, 16526A, 16192T, 16256T, 16270T Bollongino 2013
  Germany Blätterhöhle [BLA 19]   8638 ± 56 BC     U5a 16192T, 16256T, 16270T, 16294T 12308G 14798T Bollongino 2013
  England, UK Gough's Cave, Cheddar [Cheddar Man] M 8000 BC     Unreliable; U5 16192T, 16270T, reported as U5a. Never properly published. Contamination likely. Due to be re-tested. ISOGG; personal communication Chris Stringer.
  Russia Chekalino ? 7800 BC     U5a 16192T, 16256T, 16270T, 16294T, +12308 HinfI Bramanti 2009
  Russia Lebyazhinka [IV a] M 8000-7000 BC     U5a1 16192T, 16241C, 16256T, 16270T, 16399G; +12308 HinfI Bramanti 2009
  Germany Falkensteiner Höhle M 7200 BC     U5b2 12372A, 14766T, 14793A, 161721T, 167028T Bramanti 2009
  Portugal Toledo, Lourinha [K-13]   7200-6800 BC     R? 16223T Fernández 2005
  Sweden Stora Förvar cave, Stora Karlsö Island [SfF11] M 7500-7250 cal. B.P     U5a1   Skoglund 2014
  Sweden Stora Förvar cave, Stora Karlsö Island   6,873 ± 119 cal BC     U4b1   Skoglund 2013
  Germany Bad Dürrenberg [2] F 6850 BC     U4 16356C, +12308 HinfI, +7028 AluI Bramanti 2009
  Italy Grotta d’Oriente, Favignana, Ègadi Islands, Sicily [Oriente B]   8,740-8,390 cal BP     HV1? or U5b3d? 16067T. Reported as HV1, but HVR-1 only tested. 16067T also appears in U5b3d. Mannino 2012
  Germany Hohlenstein-Stadel [5830a] M 6700 BC     U5a2a 1721C, 7028T, 12372A, 14766T, 14793G, 16114A, 16192T, 16256T, 16294T, 16311C Bramanti 2009; Brandt 2013 (updated haplogroup)
  Germany Hohlenstein-Stadel [5830b] F 6700 BC     U5b2 1721T, 7028T, 12372A, 14766T, 14793A, 16192T, 16270T Bramanti 2009
Kunda Culture Lithuania Spiginas [4] F 6350 BC     U4 16356C, +12308 HinfI, +7028 AluI Bramanti 2009
Kunda Culture? Lithuania Donkalnis [1] F       U5b 16192T, 16270T, +12308 HinfI, 14793A Bramanti 2009; Brandt 2013 (updated haplogroup)
  Luxembourg Loschbour, Heffingen [LSB 1] dark hair, 50% probability of blue eyes, darker skin than Neolithic sample M 6220-5990 BC I2a1 L178+, M423+, P37.2+, L460+, M438+, L68+, P38+, M170+, M359.2-, L161.1-, L621- reported as I2a1b* U5b1a T16189C!, A6701G (reported as U5a by Delsate) Fu 2013; Delsate 2009; Lazaridis 2013; Haak 2015
Kitoi Russia Lokomotiv cemetery, Irkutsk   6125–4885 BC     A 2 samples 16223T, 16290T, 16319A Mooder 2006
Kitoi Russia Lokomotiv, Irkutsk   6125–4885 BC     C 16223T, 16298C, 16327T Mooder 2006
Kitoi Russia Lokomotiv, Irkutsk   6125–4885 BC     D 2 samples 16223T Mooder 2006
Kitoi Russia Lokomotiv, Irkutsk   6125–4885 BC     F 2 samples 16232A, 16249C, 16304C, 16311C Mooder 2006
Kitoi Russia Lokomotiv, Irkutsk   6125–4885 BC     G2a 16223T, 16227G, 16278T Mooder 2006
Kitoi Russia Lokomotiv, Irkutsk   6125–4885 BC     U5a 2 samples 16256T, 16270T Mooder 2006
  Croatia Vela Spila/Island Korčula M 6080-6020 BC     U5b2a5 16270C 16311C A16129G T16187C C16189T T16223C G16230A C16270T T16278C Szécsényi-Nagy 2014
  Poland Janisławice   6000 BC     U5b1b 16189C 16270T Stanaszek 2013
  Spain La Braña-Arintero, Leon [La Braña 2] M 7960-7750 cal BP = 5960-5750 BC     U5b2c1 As La Braña 1 Sanchez-Quinto 2012
  Spain La Braña-Arintero, Leon [La Braña 1] dark hair, probability of blue eyes, darker skin than Neolithic sample M 7940-7690 cal BP = 5940-5690 BC C1a2 M130, M216, P255, P260, V183, V199, V232, V20 Reported as C6 (nomenclature up to 24 Jan 2014) U5b2c1 150T, 723G, 960.1C. 1721T, 3197C, 6920A, 7768G, 9477A, 12372A, 13017G, 13434G, 13617C, 13637G, 14182C, 16192T, 16270T Sanchez-Quinto 2012; Olalde 2014
  Sweden Motala [1] F 5898-5531 BC     U5a1 G5460A Lazaridis 2013
  Sweden Motala [2] M 5898-5531 BC I2c2 PF3827+, L597+, P38+, U179+ , L41+, M253-, L37- U2e1 C16527T Lazaridis 2013; Haak 2015
  Sweden Motola [3] M 5898-5531 BC I2a1b* M423+, L68+, M258+, U179+, L181-, L417- U5a1 G5460A, A9389G Lazaridis 2013; Haak 2015
  Sweden Motala [4] F 5898-5531 BC     U5a2d A13158G Lazaridis 2013; Haak 2015
  Sweden Motala [6] M 5898-5531 BC I2a1* P37.2+, L55+, L232-, L159.1- U5a2d C152T!, G6480A Lazaridis 2013; Haak 2015
  Sweden Motala [9] M 5898-5531 BC I2a1a1a* L672+, P38+, P40-, Z118- U5a2 G228A, G1888A, A2246G, C3756T, G6917A, A9531G Lazaridis 2013; Haak 2015
  Sweden Motala [12] M 5898-5531 BC I2a1b L147.2+, L178+, M423+, L178+, M423+, P37.2+, L460+, L68+, M170+, M258+, U179+, M359.2-, L621- U2e1 C16527T Lazaridis 2013; Haak 2015
  Russia Sok River, Samara [SVP44/I0124] M 5650-5555 BC R1b1a M343+, L278+, [P297 equivalent PF6513+], M478-, [M478 equivalent Y13872+, Y13866- (The presence of positive and negative markers in the M478 node can reflect an intermediate stage of its formation.)], M478-, M269- U5a1d C146T, C152T, C195T, A247G, A249d, 290-291d, T489C, A769G, A825t, A1018G, A2758G, C2885T, T3552a, T3594C, G4104A, T4312C, A4715G, G7146A, C7196a, T7256C, A7521G, T8468C, A8577G, G8584A, T8655C, A9545G, C10400T, T10664C, A10688G, C10810T, C10915T, A11605t, A12217G, G13105A, A13263G, G13276A, T13506C, T13650C, T14318C, T14783C, G15043A, G15301A, A15487t, A16129G, T16187C, G16230A, T16278C, T16298C, C16311T, T16325C, C16327T, C16519T, A8577G, A11605t, A12217G, T16189C! Haak 2015; Sergey Malyshev
  Russia Yuzhnyy Oleni Ostrov [UZOO 43 and 45]   7500 BP     U2e 2 samples 16129C, 16189C, 16362C Der Sarkissian 2011; Der Sarkissian 2013
  Russia Yuzhnyy Oleni Ostrov [UZOO 16 and 40]   7500 BP     U4 2 samples 16093C, 16356C Der Sarkissian 2011; Der Sarkissian 2013
  Russia Yuzhnyy Oleni Ostrov [UZOO 70]   7500 BP     U5a 19192T, 16256T, 16270T, 16318G Der Sarkissian 2011; Der Sarkissian 2013
  Russia Yuzhnyy Oleni Ostrov [UZOO 77]   7500 BP     H? 16235G, 16311C, 16362C Der Sarkissian 2011; Der Sarkissian 2013
  Russia Yuzhnyy Oleni Ostrov [UZOO 7 and 8 ]   7500 BP     C1g Reported as C1f in Der Sarkissian 2014. 2 samples C146T, C152T, T182C!, C195T, G247A!, A249d, 290–291d, T489C, 522.AC, A769G, A825t, A1018G, A2758G, C2885T, T3552a, T3594C, G4104A, T4312C, A4715G, G7146A, C7196a, T7256C, A7521G, T8468C, A8577G, G8584A, T8655C, A9545G, C10400T, T10664C, A10688G, C10810T, C10915T, A11605t, G11914A!, A12217G, G13105A, A13263G, G13276A, T13506C, T13650C, T14318C, T14783C, G15043A, G15301A, A15487t, A16129G, A16183c, T16187C, T16189C!, G16230A, T16278C, T16298C, C16311T, T16325C, C16327T, C16519T Der Sarkissian 2011; Der Sarkissian 2013; Der Sarkissian 2014; Haak 2015
  Russia Yuzhnyy Oleni Ostrov [UZOO 74] M 5500-5000 BC R1a1* M459+, Page65.2+, M515-, M198-, M512-, M514-, L449- C1g Reported as C1f in Der Sarkissian 2014. C146T, C152T, T182C!, C195T, G247A!, A249d, 290–291d, T489C, 522.AC, A769G, A825t, A1018G, A2758G, C2885T, T3552a, T3594C, G4104A, T4312C, A4715G, G7146A, C7196a, T7256C, A7521G, T8468C, A8577G, G8584A, T8655C, A9545G, C10400T, T10664C, A10688G, C10810T, C10915T, A11605t, G11914A!, A12217G, G13105A, A13263G, G13276A, T13506C, T13650C, T14318C, T14783C, G15043A, G15301A, A15487t, A16129G, A16183c, T16187C, T16189C!, G16230A, T16278C, T16298C, C16311T, T16325C, C16327T, C16519T Der Sarkissian 2011; Der Sarkissian 2013; Der Sarkissian 2014; Haak 2015
  Russia Popova [Po 4]   7000 BP     U4 16356C Der Sarkissian 2011; Der Sarkissian 2013
  Russia Popova [Po 2]   7000 BP     U4 16093C, 16356C Der Sarkissian 2011; Der Sarkissian 2013
  Spain La Chora (Cantabria) [CH-1]   5346 ± 90 BC Date from Strauss 2002     ? 73G, 16093, 16362, reported as H6 Hervella 2012
  Spain Cingle del Mas Nou, Castellon   5000 BC     U5 2 samples 16270T Olaria 2011
  Spain Aizpea, Navarre [AIZ-1]   4600 +/- 65 BC     U5b1 16051, 16093, 16189, 16192, 1270 Hervella 2012
Narva Culture Lithuania Kretuonas [3] M 4450 BC     U5b 16192T, 16270T, +12308 HinfI, 14793A Bramanti 2009; Brandt 2013
Narva Culture Lithuania Kretuonas [1] F 4200 BC     U5b 16192T, 16270T, +12308 HinfI, 14793A Bramanti 2009; Brandt 2013
Zedmar culture? Poland Dudka [2] ? 3650 BC     U5b 16189C, 16270T, +12308 HinfI Bramanti 2009; Brandt 2013
Zedmar Culture Poland Dudka [3] ? 4000-3000 BC     U5b 16189C, 16265G, 16270T, +12308 HinfI, +7028 AluI Bramanti 2009
  Russia Serteya (Smolenskaya oblast) VIII [А3] M 4000 BC R1a1   ? 16132T, 16163T, 16166G, 16256T. Reported as H Chekunova 2014
Pitted Ware Sweden Kopingsvik, Oland [GE 33 (KOP 29)]   3200–2850 BC     T2b 16126C 16294 T 16296T 16304C Malmstrom 2015
Pitted Ware Sweden Kopingsvik [GE 44 (KOP 32)]   3200-2850 BC     H1f 16093C Malmstrom 2015
Pitted Ware Sweden Kopingsvik [GE 44 (KOP 32)]   3200-2850 BC     K1a1 16093C 16224C 16311C Malmstrom 2015
Pitted Ware Sweden Kopingsvik [GE 44 (KOP 32)]   3200-2850 BC     U4a1 16134T Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 4]   3000-2400 BC     U4 16093C 16356C Malmstrom 2015
Pitted Ware Sweden Fridtorp, Västerhejde, Gotland [Fri 24]   3000-2400 BC     U4 16093C 16356C Malmstrom 2015
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 4, 5]   3000-2400BC     U4 2 samples 16356C Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 29B]   3000-2400 BC     HV0 16298C Malmstrom 2015
Pitted Ware Sweden Visby, Gotland [Vis 30B]   3000-2400 BC     U5b1 16189C 16192 T 16270T 16362C Malmstrom 2015
Pitted Ware Sweden Fridtorp [Fri 28]   3000-2400 BC     K1a1 16093C 16224C 16311C Malmstrom 2015
Pitted Ware Sweden Visby [Vis 32]   3000-2400 BC     U5a 16192T 16256 T 16270T Malmstrom 2015
Pitted Ware Sweden Visby [Vis 7B]   3000-2400 BC     K1a1 16093C 16224C 16311C Malmstrom 2015
Pitted Ware Sweden     3000-2400 BC         Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 14, 19]   2800-2000 BC     H/R 2 samples A16129G T16187C C16189T T16223C G16230A T16278C Malmstrom 2009; Brandt 2013
Pitted Ware Sweden Ajvide [Ajv 13]   2800-2000 BC     U4 A16129G T16187C C16189T T16223C G16230A C16260T T16278C C16311T T16356C Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 52b, 66]   2800-2000 BC     U4 2 samples T16093C A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Brandt 2013
Pitted Ware Sweden Ajvide [Ajv 70] M 2800-2000 BC     U4 T16093C A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Skoglund 2012; Brandt 2013; Skoglund 2014
Pitted Ware Sweden Ajvide [Ajv 54 ]   2800-2000 BC     U5b A16129G T16187C C16189T T16223C G16230A C16270T T16278C C16311T Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 36]   2800-2000 BC     U5b2b1a A16129G T16187C C16189T T16223C G16230A C16270T T16278C C16311T T16362C Malmstrom 2009; Brandt 2013; ; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 5, 29]   2800-2000 BC     U5a1a'g 2 samples A16129G T16187C C16189T T16223C G16230A C16256T C16270T T16278C C16311T Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 52A] M 2800-2000 BC     HV0 A16129G T16187C C16189T T16223C G16230A T16278C T16298C C16311T Malmstrom 2009; Skoglund 2012; Brandt 2013; Skoglund 2014; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 53] F 2800-2000 BC     U4d   Skoglund 2014
Pitted Ware Sweden Ajvide [Ajv 58] M 2800-2000 BC I2a1 M429+, P126+, P129+, L460+, P37.2+ U4d   Skoglund 2014
Pitted Ware Sweden Ajvide [Ajv 59] M 2800-2000 BC     U   Skoglund 2014
Pitted Ware Sweden Fridtorp, Västerhejde, Gotland [Fri 15, 22]   2800-2000 BC     U4 2 samples A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Fridtorp [Fri 4]   2800-2000 BC     U5b A16129G, T16187C, C16189T, T16223C, G16230A, C16270T, T16278C, C16311T, Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Fridtorp [Fri 27]   2800-2000 BC     U5a1a'g A16129G, T16187C, C16189T, T16223C, G16230A, C16256T, C16270T, T16278C, C16311T Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 8] M 2800-2000 BC     U4d T629C A769G A825t A1018G C1224T C1226T G1315A A1811G G2174A G2182A G2185A A2758G C2885T T3594C G4104A G4153A C4189T T4312C C5211T C5974T T5999C A6047G G6075A C6181T G6755A G6993A G7146A T7256C C7412T C7427T A7521G C7974T C7978T C8351T C8375T T8468C T8655C G8701A G8860A G8998A G9192A C9540T C10088T C10213T G10398A T10664C A10688G C10810T C10873T C10915T C11332T A11337G A11467G A11914G A12308G G12372A T12705C G13105A G13276A T13506C T13650C C13860T C14620T C15049T T15693C T16093C A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Skoglund 2012; Brandt 2013; Skoglund 2014; Malmstrom 2015
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 6B]   2800-2000 BC     T2b T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T C16296T T16304C C16311T Malmstrom 2009; Brandt 2013
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 9]   2800-2000 BC     K1a T16093C A16129G T16187C C16189T T16223C T16224C G16230A C16242T T16278C Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 3]   2800-2000 BC     U4 A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Brandt 2013; ; Malmstrom 2015
  Poland Drestwo [2] M 2250 BC     U5a 16192T, 16256T, 16270T, +12308 HinfI Bramanti 2009
  Russia Bolshoy Oleni Ostrov [BOO 49-3, 57-1]   3500 BP     U4a1 2 samples 16093C, 16129A, 16311C, 16356C Der Sarkissian 2011; Der Sarkissian 2013
  Russia Bolshoy Oleni Ostrov [BOO 49-1, 72-11]   3500 BP     U5a 2 samples 16192T, 16256T, 16270T Der Sarkissian 2011; Der Sarkissian 2013
  Russia Bolshoy Oleni Ostrov [BOO 72-9, 72-10, 72-14, 78-8]   3500 BP     U5a1 4 samples 16192T, 16256T, 16270T, 16399G Der Sarkissian 2011; Der Sarkissian 2013
  Russia Bolshoy Oleni Ostrov [BOO 72-4]   3500 BP     T* 16093C, 16126C, 16294T Der Sarkissian 2011; Der Sarkissian 2013
  Russia Bolshoy Oleni Ostrov [BOO 49-2, 49-4, 57-3, 72-2, 72-7, 72-12]   3500 BP     C* 6 samples 16223T, 16298C, 16327C Der Sarkissian 2011; Der Sarkissian 2013
  Russia Bolshoy Oleni Ostrov [BOO 72-5, 72-6]   3500 BP     C5 2 samples 16148T, 16223T, 16288C, 16298C, 16311C, 16327C Der Sarkissian 2011; Der Sarkissian 2013
  Russia Bolshoy Oleni Ostrov [BOO 49-6, 72-13, 72-15]   3500 BP     D* 3 samples 16223T, 16362C Der Sarkissian 2011; Der Sarkissian 2013
  Russia Bolshoy Oleni Ostrov [BOO 49-5, 72-3]   3500 BP     Z1a 2 samples 16129A, 16185T, 16223T, 16224C, 16260T, 16298C Der Sarkissian 2011; Der Sarkissian 2013
  Russia Bolshoy Oleni Ostrov [BOO 72-1]   3500 BP     Z1a 16129A, 16155G, 16185T, 16223T, 16224C, 16260T, 16298C Der Sarkissian 2011; Der Sarkissian 2013
 

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